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When mammalian cells are exposed to formaldehyde, the levels of the reaction products of formaldehyde with the amino acids cysteine and histidine - timonacic and spinacine - are increased

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However, the UPS undergoes Kla in SLE 93 , which impacts UPS activation mediated by the metabolic switch, resulting in an inability to clear mitochondria by autophagy and a consequent increase in these organelles in mature RBCs