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doi:10.1111/j.1467-8276.2006.00948.x

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Several reports demonstrate that cigarette smoke reduces ciliary beat frequency and interrupts the intercellular coordination of the metachronal waves [18][24], and ultrastructural studies have documented smoking-associated increased incidence of structural defects in cilia, including missing radial spokes, nexin links, central sheath, outer and inner dynein arms, and central microtubules, as well as more peripheral doublets and fused cilia [25][34]