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To measure metabolites, the culture was centrifuged at 20,000 g for 3 min to remove bacterial cells, and the metabolites in the supernatant were assayed using the L-glutamate kit Yamasa NEO (Yamasa Corporation, Chiba, Japan) and F-kit (D-glucose) (Roche/R-Biopharm, Darmstadt, Germany) in accordance with kit instructions