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In contrast, itaconate, a metabolite associated with M2 macrophages, inhibits SDH, reduces the secretion of pro-inflammatory cytokines (e.g., IL-1, IL-18, IL-12, IL-6) (79), and exerts anti-inflammatory effects by blocking SDH-mediated RET and ROS generation, thereby inhibiting HIF-1 accumulation (66)

doi:10.1101/cshperspect.a011536

Li Y, Li S, Zhai Q, Hai J, Wang D, Cao M, et al

Neoadjuvant Therapy and Surgery for Advanced Thyroid Cancer

Medicine (Baltimore) 77:7382 Oh MS, Phelps KR, Traube M, Barbosa-Saldivar JL, Boxhill C, Carroll HJ (1979) D-lactic acidosis in a man with the short-bowel syndrome