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Funding: MJF was supported by T32 Funding from the University of Wisconsin Madison Medical Scientist Training Program from the National Institute of General Medical Sciences ( T32GM140935 and T32GM008692 and from the University of Wisconsin Microbes in Health and Disease from the National Institute of Allergy and Infectious Diseases ( Additional funding supporting this work was obtained from National Institute of Allergy and Infectious Diseases ( awarded to JDS as R01AI184369, R21AI173738 and R01AI137070

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