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Thanks to localization of the GGT enzyme on the outer surface of the cell membrane, -glutamyl-[1- 13 C]glycine metabolism occurs in the extracellular space and not in the cytosol or mitochondria as is the case for most HP probes including HP [1- 13 C]pyruvate 24 and [1- 13 C]dehydroascorbic acid (DHA) 34,35 , or [1,3- 13 C]acetoacetate 36,37,38 , respectively