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Sci Transl Med 12:eaax1798 Acknowledgements This work was supported by the National Key Research and Development Program of China (2022YFC2302900 and 2021YFA1300200 to CHL and JW), the National Natural Science Foundation of China (82330069, 81825014, and 31830003 to C.H.L., 82372653 to JW, and 82171744 to QC), the Shenzhen Medical Research Funding (B2302035 to CHL), the Strategic Priority Research Program of the Chinese Academy of Sciences (XDB29020000 to CHL), the State Key Laboratory of Proteomics (SKLP-K202001 to CHL), the Youth Innovation Promotion Association CAS (Y2022036 to JW), the CAS Project for Young Scientists in Basic Research (YSBR-010 to JW), and the Special Research Assistant Program of Chinese Academy of Sciences (2022000031 to QC)

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6 Biology Department/Institute of Genomics and Evolutionary Medicine, Temple University Philadelphia, PA, USA