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Glutamate dehydrogenase (GDH) and aspartate aminotransferase (AAT) catalyze the reversible reaction between glutamate and -ketoglutarate (-KG) before the incorporation of glutamate into the TCA cycle ( Figure 1A ) ( Figure 1 Compared to normal cells, cancer cells are characterized by the rapid proliferation of cells, followed by a highly active metabolism ( 1.1 Glutamate receptors and tumor microenvironment Cancer cells with different types of cells such as endothelial cells, fibroblasts, immune cells, and extracellular components ( The role of glutamate in the regulation of the metabolism in the TCA cycle has been amply investigated previously and reviewed ( Figure 1B ) [See references ( Figure 1 summarizes the different roles of glutamate in the cell)

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For urease assays, cells were diluted 1:40 in PBS and 25 l of diluted culture was spotted onto urease detection agar medium (2% urea, 1.5% agar, 0.2% monopotassium phosphate, 0.1% peptone, 0.1% dextrose, 0.5% sodium chloride, 0.0012% pH 6.8 phenol red)

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